a. State Key Laboratory of Crop Gene Exploration and Utilization in Southwest China, Sichuan Agricultural University, Chengdu, Sichuan 611130, China;
b. Triticeae Research Institute, Sichuan Agricultural University, Chengdu, Sichuan 611130, China;
c. College of Resources, Sichuan Agricultural University, Chengdu, Sichuan 611130, China
Funds:
This work was supported by the National Natural Science Foundation of China (NO. 32572313), the National Key R&D Program of China (2024YFD1201202), the Science and Technology Bureau of Sichuan Province (2023NSFSC1995 and 2025YFHZ0184), the Sichuan Science and Technology Program (2024NSFSC1968), and the Science and Technology Bureau of Chengdu City (2024-YF05-00368-SN).